Hasora chromus chromus

Butterfly Biodata:
Genus: Hasora Moore, 1881
Species: chromus Cramer, 1780
Subspecies: chromus Cramer, 1780
Wingspan of Adult Butterfly: 45mm
Caterpillar Host Plants: Pongamia pinnata (Leguminosae).

A male Common Banded Awl perching on a leaf.

A male Common Banded Awl perching on the underside of a branch.
Physical Description of Adult Butterfly:

Close-up views of the light brown and purplish scales of a female Common Banded Awl.

A male Common Banded Awl perching on the underside of a leaf.

A male Common Banded Awl perching on the underside of a grass blade.
Field Observations of Butterfly Behaviour:

A female Common Banded Awl visiting flowers.

A female Common Banded Awl perching on a stem.
Early Stages:

Host plant : Pongamia pinnata. Relatively young leaves are shown.

Two views of a `fresh' egg of the Common Banded Awl.

Two views of a maturing egg of the Common Banded Awl.

Two views of a mature egg of the Common Banded Awl with the larval head vaguely visible.

Two views of empty egg shells of the Common Banded Awl.

Two views of a 1st instar caterpillar, early in this stage, length: 1.8mm.

Two views of a 1st instar caterpillar, late in this stage, length: 2.6mm.

1st instar caterpillar, late in this stage, dormant before its moult, length: 3.2mm.

Two views of a 2nd instar caterpillar, early in this stage, length: 3.3mm.

2nd instar caterpillar, late in this stage, length: 5mm.

2nd instar caterpillar, late L2, dormant before its moult, length:5.5mm.

Two views of a 3rd instar caterpillar, early in this stage, length: 6.5mm.

Two views of a 3rd instar caterpillar, prior to its moult, length: 9mm.

Two views of a 4th instar caterpillar, early in this stage, length: 12.3mm.

Two views of a 4th instar caterpillar, late in this stage, length: 19mm.

A newly moulted 5th instar caterpillar eating its `old' larval skin.

Two views of a 5th instar caterpillar, early in this stage, length: 19mm.

Two views of a 5th instar caterpillar, length: 26mm.

Two views of a 5th instar caterpillar, late in this stage, length:36mm.

Two views of a pre-pupa of the Common Banded Awl secured via silk girdle and anchor in its shelter.

Two views of a pupa of the Common Banded Awl with a coating of whitish powdery substance.

Two views of a mature pupa of the Common Banded Awl

A newly eclosed male Common Banded Awl.
References:

Butterfly Biodata:
Genus: Hasora Moore, 1881
Species: chromus Cramer, 1780
Subspecies: chromus Cramer, 1780
Wingspan of Adult Butterfly: 45mm
Caterpillar Host Plants: Pongamia pinnata (Leguminosae).

A male Common Banded Awl perching on a leaf.

A male Common Banded Awl perching on the underside of a branch.
Physical Description of Adult Butterfly:
Adults are rather large in size with pointed forewing apex and markedly lobate hindwings. Above,  the wings are dark brown. In the male,  the wings are unmarked while  the female has two pale yellowish crescentic spots in spaces 2 and 3 on  the forewing. The male has a discal stigma on the forewing.  Below,  both sexes are pale purplish brown.  On the hindwing, there is a  prominent and narrow  white to bluish white discal  band partitioning  the wing into an inner and an outer half. The inner edge of the band is  more sharply defined compared to the outer edge.  In contrast, the  discal band on the forewing is faint and extends only over the upper  portion of the wing. The female has two apical spots in spaces 6 and 7  of  the forewing with the one in space 7 absent from some specimens.

Close-up views of the light brown and purplish scales of a female Common Banded Awl.

A male Common Banded Awl perching on the underside of a leaf.

A male Common Banded Awl perching on the underside of a grass blade.
Field Observations of Butterfly Behaviour:
This species is  uncommon in Singapore. There is a better chance of finding larval stages of this species on its host plant, Pongamia pinnata,  than seeing the adults up close. The fast flying adults have been  observed to visit flowers for nectar.  When resting, the adults  typically do so on the underside of a leaf or other plant parts. 

A female Common Banded Awl visiting flowers.

A female Common Banded Awl perching on a stem.
Early Stages:
The local host plant, Pongamia pinnata,  is a tree which can grow up to 6-15m tall. Its compound leaf has 5-7  leaflets in a pinna. Each leaflet is thin leathery, shiny reddish brown  when young, and turn darker green when matured. It is listed as  "Endangered" in the latest Singapore Red Data book, and at present can  be found locally in small numbers in  parks and along side walks in  certain built-up areas. The caterpillars of the Common Banded Awl feed  on young to middle-aged leaves of this plant.

Host plant : Pongamia pinnata. Relatively young leaves are shown.
The  eggs are laid singly or in small groups of 2-3  on young shoots of the  host plants. Each egg is initially white and shaped like a bun with a  flattened base (diameter: 0.4-0.5mm).  Prominent ridges running from the  pole to the base. The micropylar sits atop at the pole. The entire egg  turns pinky red as it develops, and then decolorizes again when the  caterpillar is ready to emerge.

Two views of a `fresh' egg of the Common Banded Awl.

Two views of a maturing egg of the Common Banded Awl.

Two views of a mature egg of the Common Banded Awl with the larval head vaguely visible.

Two views of empty egg shells of the Common Banded Awl.
It  takes 2-3 days for the collected egg to hatch. The young caterpillar  eats just enough of the shell to emerge, and has a length of about 2mm.  It has the typical cylindrical shape for skipper caterpillars, and the  yellowish brown body has a number of short setae. The large head is  black, slightly bi-lobed and lightly hairy. The dorsum of the prothorax  carries a dark-colored patch/shield.  

Two views of a 1st instar caterpillar, early in this stage, length: 1.8mm.

Two views of a 1st instar caterpillar, late in this stage, length: 2.6mm.
The  young caterpillar constructs its first leaf shelter by bringing  two  halves of a small young leaf together  with silk threads. It rests  within the flap and ventures out to eat on nearby leaf surface. In  later instars, the Common Banded Awl caterpillars also construct leaf  shelters in a similar fashion but do so with larger and older leaves.   The larval  growth of the Common Banded Awl is rapid in pace for  all  five of its instars, with   all but the final  instar lasting only 1-1.5  days.

1st instar caterpillar, late in this stage, dormant before its moult, length: 3.2mm.
After  reaching a length of about 3.2mm in the 1st instar, the caterpillar  moults to the next instar after a short dormant period. The 2nd instar  caterpillar has four faint  whitish narrow dorsal bands  and one lateral  band which are barely visible against the pale yellowish body ground  color.  These bands become more prominent as the body ground colour turn  pale brownish towards the end of this instar.  The body and the head  capsule are also covered in short fine setae.

Two views of a 2nd instar caterpillar, early in this stage, length: 3.3mm.

2nd instar caterpillar, late in this stage, length: 5mm.

2nd instar caterpillar, late L2, dormant before its moult, length:5.5mm.
The  2nd instar caterpillar reaches a length of about 5.5mm, and after 2  days in this stage, it moults again. The 3rd instar caterpillar  resembles the 2nd instar caterpillar but with more prominent  dorsal and  lateral whitish bands due to the much darker brown in its ground  colour.  Overall the body takes on a freckled appearance with numerous  tiny lighter patches dotting the dark brown surface. A narrow anal   plate  dark brown to black in color, can now be seen on the posterior  end. This instar lasts another  1-1.5  days with the length reaching  9mm. 

Two views of a 3rd instar caterpillar, early in this stage, length: 6.5mm.

Two views of a 3rd instar caterpillar, prior to its moult, length: 9mm.
The  4th instar caterpillar resembles the 3rd instar caterpillar closely,  but with its body ground colour in  darker brown. The whitish setae on  the head capsule are  proportionately much longer than in earlier  instars. This stage takes about 1-1.5 days to complete with the body  length reaching 19mm. 

Two views of a 4th instar caterpillar, early in this stage, length: 12.3mm.

Two views of a 4th instar caterpillar, late in this stage, length: 19mm.
The  final and 5th instar caterpillar has similar body markings as the 4th  instar caterpillar, though the ground colour has changed to a dark shade  of purplish brown or even pinky brown above the lateral band. Several  large dorso-lateral spots are also present here. Below the narrow  lateral band, the body is in a contrasting yellowish brown. One  unmistaken change is in the head capsule which has now become  pale  orangy brown.  Moderately long white setae are found  over the whole  body and the head capsule. This stage takes about 3 days to complete  with body length reaching up to  36mm.

A newly moulted 5th instar caterpillar eating its `old' larval skin.

Two views of a 5th instar caterpillar, early in this stage, length: 19mm.

Two views of a 5th instar caterpillar, length: 26mm.
Towards  the end of 5th instar, the body of the caterpillar gradually shrinks in  length and decolorises. Some caterpillars would turn completely pale  greenish whilst others retains several lateral dark spots. The fully  grown caterpillar ceases feeding and  stations itself in its leaf  shelter and enters the preparatory pupa phase. During the early part of  this stage, the caterpillar spins large quantity of silk threads to seal  the pupation shelter, and in particular, constructs a silk girdle at  its 2nd/3rd abdominal segment and a short transverse silk band near its  posterior end. Both the dorsal point of the girdle and the transverse  band are also secured by vertical/oblique threads to the inner wall of  the shelter.

Two views of a 5th instar caterpillar, late in this stage, length:36mm.

Two views of a pre-pupa of the Common Banded Awl secured via silk girdle and anchor in its shelter.
After  about 0.5-1 day of the pre-pupal phase,  pupation takes place within  the  pupation shelter. The pupa secures itself with its cremaster  attached to the  transverse band. The pupa has a short thorax, a rather  long abdomen and a short and pointed rostrum. Fresh after the pupation  event,  the body is greenish, but after 0.5 to 1 day, the body surface  becomes mostly covered in a white substance.  Length of pupae: 23-24mm.

Two views of a pupa of the Common Banded Awl with a coating of whitish powdery substance.
After  5-5.5 days, the pupa becomes darkened in color signaling the imminent  emergence of the adult. The next day the adult butterfly emerges from  the mature pupa.

Two views of a mature pupa of the Common Banded Awl

A newly eclosed male Common Banded Awl.
References:
- The Butterflies of The Malay Peninsula, A.S. Corbet and H.M. Pendlebury, 4th Edition, The Malayan Nature Society.
 - Butterflies of Thailand, Pisuth Ek-Amnuay, 1st Edition, 2006
 - The Butterflies of Hong Kong, M. Bascombe, G. Johnston, F. Bascombe, Princeton University Pres 1999
 
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